Courses I have recently taught, am teaching or will teach at the University of Guelph
BIOL*3040 Methods in
This course will
provide students with an understanding of the major analytical approaches used in modern evolutionary biology and
an appreciation of the relevance of these methods to other branches of the life sciences. This includes the
analysis of molecular data, phylogenetics and “tree thinking”, population genetics, genomics, phenotypic selection,
experimental evolution, and hypothesis generation and testing in historical sciences. Laboratory sessions will be
devoted to practical training in analytical tools using specialized computer software and real datasets. Students
will also undertake an independent project in order to put these approaches into action.
This course provides a broad overview of
evolutionary biology. It examines the concepts and mechanisms that explain evolutionary change and the evolution of
biological diversity at different levels of biological organization (gene to ecosystem) and across space and time.
It also introduces historical forms of scientific inquiry, unique to biology. The course is designed to be of
interest to students with general interests in science and in research in all areas of biology.
BIOL*3020 Population Genetics
This course is designed to explore the
concepts of random mating, inbreeding, random drift, assortative mating and selection as they relate to natural
populations. The dynamic genetic structure of populations and its relationship to the process of speciation is
examined. The role and significance of molecular genetics as it relates to population genetics, evolution,
systematics and phylogeny is also considered. Department of Integrative Biology.
ZOOL*3700 Integrative Biology of
This course explores variation in
physiology, reproduction and life history among invertebrates, and the role of invertebrates in marine, freshwater
and terrestrial ecosystems. Through field experiences, lab study and a class experiment, students will study the
diverse functions including: circulation and gas exchange; feeding and digestion; osmoregulation and
excretion, nervous system and sensory structures and locomotion, that allow invertebrates to live in very different
I generally supervise one or
students per year in 4th-year undergraduate research
IBIO*4521 and IBIO*4522 Undergraduate Thesis in Integrative
This two-semester course series involves
independent research of a practical or theoretical nature on a specific topic in biology. Research is carried
out under the supervision of an individual faculty member. Students must write and defend a thesis based on the
results of their work at the end of semester 2. Arrangements must be made with both a faculty supervisor and
the course coordinator at least one semester in advance of taking the course.
IBIO*4500 and IBIO*4510 Research Project in Integrative
These one-semester courses involve
independent research of a practical or theoretical nature on a specific topic in biology. Research is
carried out under the supervision of an individual faculty member. Students should make arrangements with both a
faculty supervisor and the course coordinator at least one semester in advance of taking the course.
MCB*4500 and MCB*4510 Research Project in Molecular and Cellular
This two-semester course series involves
independent research of a practical or theoretical nature on a specific topic in molecular and cellular
biology. Research is carried out under the supervision of an individual faculty member. Arrangements should be
made with both a faculty supervisor and the course coordinator at least one semester in advance of taking the
Advances in Evolutionary Biology
This modular graduate course reviews
books and/or other publications in the field of evolutionary biology, providing knowledge of progress in this area
of biology. Topics may include epigenetics, phylogenetics, developmental basis of evolutionary change, and
molecular evolution. The course includes lectures and seminars in which the students participate.
The course was based on the
2nd edition of Graham Bell’s book
Selection: the mechanism of