Faculty | MSc| PhD| Shared | Courses
Chair - Paul D.N. Hebert (359 Axelrod, Ext. 3598/8763)
Graduate co-ordinator - James S. Ballantyne (379 Axelrod, Ext. 2708/8385)
Graduate secretary - Mary Anne Davis (255 Axelrod, Ext. 6094)
James S. Ballantyne BSc, MSc Guelph, PhD British Columbia - Associate
F. William H. Beamish BA, PhD Toronto - Professor
Jim P. Bogart BSA Toronto, MA, PhD Texas - Professor
Elizabeth G. Boulding BSc British Columbia, MSc Alberta, PhD Washington -
Ronald J. Brooks BSc, MSc Toronto, PhD Illinois - Professor
George A. Bubenik MUDr Charles - Associate Professor
Teresa J.D. Crease BSc, MSc Windsor, PhD Washington - Assistant
Roy G. Danzmann BSc, MSc Guelph, PhD Montana - Assistant Professor
Moira M. Ferguson BSc, MSc Guelph, PhD Montana - Associate
John M. Fryxell BSc, PhD British Columbia - Associate Professor
David E. Gaskin BSc Bristol, PhD Massey - Professor
Paul D.N. Hebert BSc Queen's, PhD Cambridge, FRSC - Professor
Denis H. Lynn BSc Guelph, PhD Toronto - Professor
Gerald L. Mackie BSc, MSc, PhD Ottawa - Professor
Alexander L.A. Middleton BSc, MSc Western Ontario, PhD Monash -
David L.G. Noakes BSc, MSc Western Ontario, PhD California -
Thomas D. Nudds BSc, MSc Windsor, PhD Western Ontario - Professor
John C. Roff BA, MA Cambridge, PhD Newcastle - Professor
Michael E. Ruse BA, PhD Bristol, MA McMaster, Dr. Philos, H.C. (Bergen),
FRSC - Professor
Steven R. Scadding BSc, MSc, PhD Toronto - Associate Professor
E. Donald Stevens BSc, MSc, PhD British Columbia - Professor
Vernon G. Thomas BA Oxford, MSc, PhD Guelph - Associate Professor
Glen J. Van Der Kraak BSc, MSc Manitoba, PhD British Columbia - Associate
Patrick T.K. Woo BSc, MSc British Columbia, PhD Guelph - Professor
Patricia A. Wright BSc McMaster, PhD British Columbia - Assistant
Peter Yodzis BSc Duke, PhD New Mexico State - Professor
Associated Graduate Faculty
Roy C. Anderson BSc Alberta, MA, PhD Toronto, Dip. Helm Paris - University
Eugene K. Balon RNDr, CSc Charles - University Professor Emeritus
D. George Dixon Bsc Sir George Williams, MSc Concordia, PhD Guelph -
University of Waterloo
Robert J. Etches BSc (Agr) British Columbia, MSc McGill, PhD, DSc Reading
- Adjunct Professor
John C. George BSc, PhD Bombay - University Professor Emeritus
Arthur H. Houston BSc McMaster, MA, PhD British Columbia - Adjunct
Professor, Brock University
Sarah A. Jones BSc Nottingham, PhD Bristol - Mount Sinai Hospital Research
John S. Millar BA, MA British Columbia, PhD Alberta - University of Western
Derek C.G. Muir BSc, MSc, PhD McGill - Department of Fisheries and Oceans,
Kelly R. Munkittrick BSc, MSc Guelph, PhD Waterloo - Department of
Fisheries and Oceans, Burlington
Mark S. Ridgway BSc Miami, MSc British Columbia, PhD Western Ontario -
Ontario Ministry of Natural Resources
John B. Sprague BSc, MSc, PhD British Columbia - Retired
William D. Taylor BSc, PhD Toronto - University of Waterloo
John H. Youson BA Victoria, MSc McGill, PhD Western Ontario - University
Special Graduate Faculty
McLaughlin, Robert L. BSc Queen's, MSc Windsor, PhD McGill - Research
G. Fred Ramprashad BSc Western Ontario, MSc Guelph - Associate Professor
Watson, Alan P. BSc, MSc Guelph - Director, Arboretum
The Department of Zoology comprises graduate students and
faculty with diverse research interests. These interests are concentrated in three major areas of
emphasis: ecology and behaviour, evolutionary biology, and physiology.
The Department of Zoology offers MSc degrees in each of the three major
areas of emphasis, focusing on (but not restricted to) experimental approaches in field and
laboratory settings and a strong linkage between theoretical and applied investigations. The
department encourages students to pursue interdisciplinary research and, where appropriate,
utilize faculty expertise from across campus on their advisory committees.
To be considered, applicants must meet the requirements of a four-year
honours science degree with a minimum 'B' (73%) average during the final two years (4
semesters) of undergraduate study. Applicants must obtain the support of a faculty member
willing to serve as their thesis adviser. For more information regarding this requirement,
applicants should consult the section titled "How to Apply to Graduate School" in the
department's booklet Research in Zoology, available from the office of the graduate
secretary of Zoology.
Under exceptional circumstances, an applicant with a 'B-' (70-72%)
average during the last 2 years (four semesters) of study may be considered for admission. Such
applicants must have outstanding letters of recommendation that provide strong evidence of
potential research capability and a strong endorsement from a potential thesis adviser.
Admission may be granted in September, January or May. Completed
applications should arrive in the department at least one full semester (four months) before the
expected date of admission. Applications from international students, especially those applying
for financial support, should arrive at least eight months prior to the expected date of
Students must complete and defend an acceptable thesis. In addition, they
must successfully complete courses totalling not fewer than 1.5 credits.
An acceptable MSc thesis comprises a scientifically defensible account of
the student's research on a particular, well-defined research problem or hypothesis. (Such
research should begin with the practical expectation that it could be completed and the thesis
defended in not more than six semesters.) Paramount to the notion of acceptability of the thesis is
its quality with respect to the underlying rationale (problem identification), the approach used to
address the problem, and the evaluation of the results. Final acceptance of the MSc thesis need
not imply that the work is sufficiently meritorious to warrant publication in scholarly media,
though the majority of MSc research in the department is published.
The department endorses the idea that graduate students in zoology should
benefit from exposure to recent developments both within and between the major areas of
emphasis. To that end, students may enrol in any of the regularly offered courses entitled
"Advances in ...", which are team-taught by several faculty members . A selection of subjects is
given in each of the course descriptions below. Details of course content, format and evaluation
will be available in the office of the chair of the department one semester prior to the semester in
which the course is offered. In addition, the department offers two "Topics in..." courses to
provide students with the opportunity to study with individual faculty on specific topics in the
faculty member's area of expertise. These courses may be taken by groups as either
reading/seminar courses, or on an individual research-project basis. Students should approach
individual faculty members to request supervision on individual research project courses; faculty
members may be petitioned by students to offer, or may advertise, "Topics in..." courses at least
one semester prior to the semester in which the course is to be offered.
The Department of Zoology offers PhD degrees for studies in each of the
three major areas of emphasis: ecology and behaviour, evolutionary biology, and
The admission and degree requirements of the PhD program are essentially
those of the university. Most applicants will have a recognized master's degree in a related field
obtained with minimum academic standing of 'A-' (80%) in their postgraduate studies, and the
endorsement of a potential thesis adviser. For more information about this last requirement,
applicants should refer to the department's booklet Research in Zoology, available from
the graduate secretary of Zoology. Under exceptional circumstances admission directly to a PhD
program with an appropriate honours degree alone, or transfer from MSc to PhD program
without completing the MSc thesis requirements, is also possible. Applications should be
received at least one full semester (four months) prior to the expected date of admission.
Applications from international students, especially those applying for financial support, should
arrive at least eight months prior to the expected date of admission.
The Department of Zoology expects that the major part of the student's
time will be devoted to research in fulfilment of the thesis requirement. For that reason, the
department does not require that PhD students take any courses. Even so, advisory committees
may, from time to time, require that a student take some prescribed or additional courses.
Regardless, PhD students are expected to contribute and participate actively in the full academic
life of the department, including regular attendance at departmental and inter-departmental
seminars, and to provide leadership and counselling to undergraduate and MSc students.
PhD students will become candidates for the PhD degree upon
successful completion of an oral qualifying examination, which must be conducted not later than
the fifth semester of the PhD program. The exam evaluates students' knowledge in the general
area of the intended research.
Candidates will spend not fewer than five semesters (seven without an
MSc) in the program, and are expected to complete their studies within 11 semesters.
Submission and defence of an acceptable thesis complete the requirements
for a PhD. An acceptable thesis comprises a report of the candidate's research on a particular and
well-defined research problem or hypothesis. It should represent a significant contribution to
knowledge in that field. Emphasis is placed on the quality of the work as judged by the
expression of mature scholarship, critical judgment, and satisfactory literary style in the thesis.
Thesis approval implies that it is judged sufficiently meritorious to warrant publication in
reputable, refereed journals in its field.
Msc (Aquaculture) Interdepartmental Program
The Department of Zoology participates in the MSc program in
aquaculture. Those faculty members whose research and teaching expertise includes aspects of
aquaculture may serve as advisers for MSc (Aquaculture) students. Please consult the
Aquaculture listing for a detailed description of the MSc (Aquaculture) interdepartmental
Biophysics MSc/PhD Program
The Department of Zoology participates in the MSc/PhD program in
biophysics. Those faculty members whose research and teaching expertise includes aspects of
biophysics may serve as advisers for MSc and PhD students in biophysics. Please consult the
Biophysics listing for a detailed description of the graduate programs offered by the Biophysics
Interdepartmental Group (BIG).
Toxicology MSc/PhD Collaborative Program
The Department of Zoology participates in the MSc/PhD program in
toxicology. Those faculty members whose research and teaching expertise includes aspects of
toxicology may serve as advisers for MSc and PhD students. Please consult the Toxicology
listing for a detailed description of the MSc/PhD collaborative program.
- 9206020 Advances in Evolutionary Biology (0.5)
- This modular course reviews books and/or other publications in the field of
evolutionary biology, providing knowledge of progress in this area of biology. Topics may
include epigenetics, phylogenetics, developmental basis of evolutionary change, and molecular
evolution. The course includes lectures and seminars in which the students participate. Offered
- 9206010 Advances in Physiology (0.5)
- A modular course format in which several faculty members lecture and/or lead
discussion groups in tutorials on advances in their areas, or related areas, of physiology. Topics
may include metabolic adaptation to extreme environments, behavioural and molecular
endocrinology, and exercise and muscle physiology. The course includes lectures and seminars in
which the students participate. Offered annually.
Ecology and Behaviour
- 9206000 Advances in Ecology and Behaviour (0.5)
- This is a modular course in which several faculty lecture and/or lead discussion
groups in tutorials about advances in their broad areas, or related areas, of ecology and
behaviour. Topics may include animal communication, optimal foraging, life-history evolution,
mating systems, population dynamics, niche theory and food-web dynamics. The course includes
lectures and seminars in which the students participate. Offered annually.
- 9206070 Topics in Advanced Zoology I (0.5)
- 9206080 Topics in Advanced Zoology II (0.5)
- These courses provide graduate students, either individually or in groups, with the
opportunity to pursue topics in specialized fields of zoology under the guidance of graduate
faculty. Course topics will normally be advertised by faculty one semester prior to their offering.
Courses may be offered in any of lecture, reading/seminar, or individual project formats. A
minimum enrolment may be required for some course offerings.
- 9206550 Aquaculture (0.5)
- Examination of the history, practice and future of aquaculture with special
reference to the application of biological principles and knowledge to the production of aquatic
organisms for food and other uses.
- 9206630 Seminar in Scientific Communication (0.5)
- The development and refinement of the skills of communication (oral and
- 9206640 Research Seminars in Zoology (0.25)
- Participation in a regular departmental seminar series including a presentation.