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Dr. Cortland Griswold
Associate Professor

Dr. Cortlant Griswold


Office: SSC 1474
Ext: 56240
Lab: SSC 1409/1410
Ext: 56014

The Griswold Theoretical Biology Group


I am an evolutionary biologist whose research focuses on theoretical and inference questions.  My interest in evolutionary theory and inference began the second term of my third year as an undergraduate.  I attended a series of lectures that introduced the theory of population genetics and demonstrated how ecological processes bring about changes in allele frequencies, i.e. genetic evolution by natural selection.  At the time I had very broad interests in biology, participating in research on vocal communication in birds and ecological physiology in plants.  I continued pursuing my interest in understanding the evolution of bird vocalizations as an M.Sc. student and this work heightened again my interest in using theory and population genetics principles to help us understand ourselves and the natural world around us.  I then pursued a Ph.D. in the area of theoretical population and evolutionary genetics.


B.Sc. – University of Wisconsin-Madison
M.Sc. – University of Toronto
Ph.D. – University of British Columbia


Our group seeks to advance population genetic and evolutionary theory and to develop computational methods that address basic and applied questions in biology. In particular, a focus of my research program is to characterize processes that facilitate the evolution of multivariate diversity. Graduate student research in my lab is not limited to my research program and is in fact quite diverse; for more information, see my lab's webpage.

Selected Publications

Betini, G.S., C.K. Griswold, and D.R. Norris. 2013. Density-mediated carry-over effects explain variation in breeding output across time in a seasonal population. 9: 20130582.

Betini, G.S., C.K. Griswold, and D.R. Norris. 2013. Carry-over effects, sequential density dependence and the dynamics of populations in a seasonal environment. Proceedings of the Royal Society B 280: 20130110.

Griswold, C.K. and T.A. Henry. 2012. Epistasis can increase multivariate trait diversity in haploid non-recombining populations. Theoretical Population Biology 82: 209-221.

Griswold, C.K. and D.J. Eisner. 2012. The mapping of epistatic effects onto a genealogical tree in haploid populations. Theoretical Population Biology 81: 32-44.

Griswold, C.K., C.M. Taylor and D.R. Norris. 2011. The equilibrium population size of a partially migratory population and its response to environmental change. Oikos 120: 1847-1859.

Griswold, C.K. 2011. A Model of the Physiological Basis of a Multivariate Phenotype that is Mediated by Ca2+ Signaling and Controlled by Ryanodine Receptor Composition. Journal of Theoretical Biology 282: 14-22.

Griswold, C.K., C.M. Taylor and D.R. Norris. 2010. The evolution of migration in a seasonal environment. Proceedings of the Royal Society B 277: 2711-2720.


BIOL*2400 Evolution
BIOL*3020 Population Genetics