Cortland Griswold

Associate Professor
Phone number: 
519-824-4120 x56240
SSC 1474


  • B.Sc. – University of Wisconsin-Madison
  • M.Sc. – University of Toronto
  • Ph.D. – University of British Columbia


Broadly my research group's interests are in theoretical biology, with a focus on evolutionary genetics and evolutionary theory.

Current projects in my research group are

[1] Developing ancestral graph methods to understand the evolution of multivariate diversity

[2] Multilocus selection in polyploids

[3] Plasmid population genetics

[4] Metagenomic theory

Students with interests in theoretical biology are welcome to contact me about undergraduate or graduate research.

Select Publications

  • Griswold, C.K. and M.W. Williamson. 2017.  A two-locus model of selection in autotetraploids: Chromosomal gametic disequilibrium and selection for an adaptive epistatic gene combination.  Heredity 119: 314-327.
  • Betini, G.S., A. McAdam, C.K. Griswold and D.R. Norris.  2017.  A fitness trade-off between seasons causes multigenerational cycles in phenotype and population size. eLife 6: e18770.
  • Griswold, C.K.  2015.  Additive genetic variation and evolvability of a multivariate trait can be increased by epistatic gene action.  Journal of Theoretical Biology 387: 241-257.
  • Griswold, C.K.  2015.  Epistasis can accelerate adaptive diversification in haploid asexual populations.  Proceedings of the Royal Society B 282: 20142648.
  • Betini, G.S., A. Pardy, C.K. Griswold, and D.R. Norris. 2015. Causes and consequences of density-dependent dispersal: the role of seasonality and carry-over effects. Ecosphere 6: 1-9.
  • House, M.A., C.K. Griswold and L. Lukens. 2014.  Evidence for selection on gene expression in cultivated rice (Oryza sativa).  Molecular Biology & Evolution 31: 1414-1525.
  • Betini, G.S., C.K. Griswold, L. Prodan and D.R. Norris.  2014.  Body size, carry-over effects and survival in a seasonal environment: consequences for population dynamics.  Journal of Animal Ecology 83: 1313-1321.
  • McKay, P.B. and C.K. Griswold.  2014. A comparative study indicates both positive and purifying selection within ryanodine receptor (RyR) genes, as well as correlated evolution.  Journal of Experimental Zoology Part A: Ecological Genetics & Physiology 321: 151-163.
  • Betini, G.S., C.K. Griswold, and D.R. Norris.  2013.  Density-mediated carry-over effects explain variation in breeding output across time in a seasonal population.  Biology Letters 9: 20130582.
  • Betini, G.S., C.K. Griswold, and D.R. Norris.  2013.  Carry-over effects, sequential density dependence and the dynamics of populations in a seasonal environment.  Proceedings of the Royal Society B 280: 20130110.
  • Griswold, C.K. and T.A. Henry.  2012.  Epistasis can increase multivariate trait diversity in haploid non-recombining populations.  Theoretical Population Biology 82: 209-221.
  • Griswold, C.K. and D.J. Eisner.  2012.  The mapping of epistatic effects onto a genealogical tree in haploid populations.  Theoretical Population Biology 81: 32-44.
  • Griswold, C.K. 2011. A Model of the Physiological Basis of a Multivariate Phenotype that is Mediated by Ca2+ Signaling and Controlled by Ryanodine Receptor Composition. Journal of Theoretical Biology 282: 14-22.
  • Griswold, C.K., C.M. Taylor and D.R. Norris. 2011. The equilibrium population size of a partially migratory population and its response to environmental change.  Oikos 120: 1847-1859.


  • BIOL*3020 Population Genetics
  • BIOL*3040 Methods in Evolutionary Biology
  • IBIO*6000 Foundations in Theoretical Biology